1. Department of Biology, University of York, Heslington YO10 5DD, UK
2. Institute of Molecular Cell and Systems Biology, College of Medical Veterinary and Life Sciences, University of Glasgow, Glasgow G12 8QQ, UK
3. Institute of Cardiovascular and Medical Sciences, College of Medical Veterinary and Life Sciences, University of Glasgow, Glasgow G12 8QQ, UK
4. School of Medicine, Dentistry & Nursing, College of Medical Veterinary and Life Sciences, University of Glasgow, Glasgow G12 8QQ, UK
5. Strathclyde Institute of Pharmacy and Biomedical Sciences, University of Strathclyde, 161 Cathedral Street, Glasgow G4 ORE, UK