1. Bodega Marine Laboratory, University of California, Davis, 2099 Westshore Road, Bodega Bay, California 94923, Graduate Group in Ecology, University of California, Davis, California 94923;, Email: gha@ucdavis.edu
2. Bodega Marine Laboratory, University of California, Davis, 2099 Westshore Road, Bodega Bay, California 94923; Department of Evolution and Ecology, University of California, Davis, One Shields Avenue, Davis, California 95616