1. Department of Medicine and Bioregulatory Science, Graduate School of Medical Sciences, Kyushu University , Fukuoka, 812-8582 , Japan
2. Institute for Integrated Cell-Material Sciences (WPI-iCeMS), Kyoto University , Kyoto, 606-8501 , Japan
3. Center for Animal Disease Models, Research Institute for Biomedical Sciences, Tokyo University of Science , Yamazaki 2669 , Noda-shi, Chiba, 278-0022, Japan