1. College of Fishery Guangdong Ocean University Zhanjiang524088 China
2. Guangdong Provincial Key Laboratory of Pathogenic Biology and Epidemiology for Aquatic Animals Zhanjiang 524088China
3. Guangdong Key Laboratory of Control for Diseases of Aquatic Economic Animals Zhanjiang 524088China
4. Department of Fisheries and Aquatic Resources Management University for Development Studies Tamale‐Ghana China
5. Langye Animal Husbandry Company Limited Gaozhou City Guangdong Province China