1. Instituto Gulbenkian de Ciência; Oeiras Portugal
2. Faculté des Sciences; Université de Mahajanga; Mahajanga Madagascar
3. Centre for Research into Ecological and Environmental Modelling; The Observatory, University of St. Andrews; St. Andrews Scotland United Kingdom
4. Laboratoire Evolution and Diversité Biologique, CNRS; Université Paul Sabatier, ENFA; Toulouse France
5. Université de Toulouse; Toulouse France