1. Population Health Department; QIMR Berghofer Medical Research Institute, 300 Herston Road, Herston; QLD Australia
2. Faculty of Medicine, Herston Road; The University of Queensland; Herston QLD Australia
3. Cancer Research UK Manchester Institute and Institute of Inflammation and Repair, University of Manchester; Manchester United Kingdom
4. Department of Gastroenterology and Hepatology; Princess Alexandra Hospital; Brisbane QLD Australia