1. Department of Microbiology and Immunology University of Melbourne at the Peter Doherty Institute for Infection and Immunity Melbourne VIC 3000 Australia
2. Australian Research Council Centre of Excellence in Advanced Molecular Imaging University of Melbourne Parkville VIC 3010 Australia
3. School of BioSciences University of Melbourne Parkville VIC 3010 Australia
4. John Curtin School of Medical Research The Australian National University Canberra ACT 2600 Australia