1. Gastrointestinal Stem Cell Biology Laboratory, Wellcome Trust Centre for Human Genetics; University of Oxford; Roosevelt Drive Oxford UK
2. Translational Gastroenterology Unit, Experimental Medicine Division, Nuffield Department of Clinical Medicine; John Radcliffe Hospital; Headington Oxford UK
3. Molecular and Population Genetics Laboratory, Wellcome Trust Centre for Human Genetics; University of Oxford; Roosevelt Drive Oxford UK
4. South Australian Health and Medical Research Institute; University of Adelaide; Adelaide South Australia Australia