1. U.S. Geological Survey; Wisconsin Cooperative Wildlife Research Unit; University of Wisconsin; Madison Wisconsin 53706 USA
2. U.S. Geological Survey; Pacific Island Ecosystems Research Center; Hawaiʻi National Park; Hawaiʻi 96718 USA
3. University of New England; Biddeford Maine 04005 USA
4. University of Hawaiʻi at Hilo; Hilo Hawaiʻi 96720 USA