1. Department of Biology, Geomatics and Landscape Ecology Laboratory Carleton University Ottawa Ontario Canada
2. Canadian Wildlife Service Environment and Climate Change Canada Yellowknife Northwest Territories Canada
3. Centre d'Écologie Fonctionnelle et Évolutive CNRS Montpellier Cedex 5 France
4. UMR Eco&Sols INRAE Montpellier Cedex 2 France
5. Wildlife Research Division Environment and Climate Change Canada Ottawa Ontario Canada