1. Institut de Systématique, Évolution, Biodiversité (ISYEB) Muséum National d'Histoire Naturelle Sorbonne Université CNRS EPHE; CP 30, 25 rue Cuvier 75005 Paris, France
2. Museum of Paleontology & Department of Earth & Environmental Sciences University of Michigan 1105 N University Ave Ann Arbor MI 48109‐1079 USA
3. School of Geography Earth & Environmental Sciences University of Birmingham Birmingham B15 2TT UK
4. Christ Church Oxford OX1 1DP UK