1. Sea Mammal Research Unit Scottish Oceans Institute University of St Andrews St Andrews KY16 8LB United Kingdom
2. Centre for Environment, Fisheries and Aquaculture Science Pakefield Road Lowestoft NR33 0HT United Kingdom
3. Centre for Research into Ecological and Environmental Modelling The Observatory University of St Andrews St Andrews KY16 9LZ United Kingdom
4. Lighthouse Field Station Institute of Biological and Environmental Sciences University of Aberdeen George Street Cromarty IV11 8YL United Kingdom