1. Ecologie Systématique EvolutionUniv. Paris‐Sud CNRS AgroParisTech, Université Paris‐Saclay 91400 Orsay France
2. ISA UMR INRA CNRS Universite Nicé Côte d'Azur 06903 Sophia‐Antipolis France
3. CBGP UMR INRA CIRAD IRD SupAgro 34988 Montferrier sur Lez France
4. Department of Ecology and Evolutionary BiologyCenter for Tropical ResearchInstitute of the Environment and SustainabilityUniversity of California Los Angeles Los Angeles California 90095 USA