1. Biological Resource Management Division; National Park Service; 1201 Oakridge Drive 200 Fort Collins CO 80525 USA
2. Colorado State University; Veterinary Diagnostic Laboratory; 300 W Drake Road Fort Collins CO 80523 USA
3. United States Department of Agriculture; Animal and Plant Health Inspection Service; National Wildlife Research Center; 4101 LaPorte Avenue Fort Collins CO 80525 USA
4. Animal Reproduction and Biotechnology Laboratory; Colorado State University; Fort Collins CO 80523 USA