1. Université Grenoble Alpes; F-38000 Grenoble France
2. CEA, LETI; MINATEC Campus F-38054 Grenoble France
3. Laboratory of Plant & Cell Physiology; CEA/DRF/BIG; CNRS UMR5168, INRA UMR 1417 F-38054 Grenoble France
4. INAC/SyMMES; UMR 5819 CEA CNRS UGA; MINATEC Campus F-38054 Grenoble France