1. Institute of Geosystems and Bioindication Technische Universität Braunschweig Braunschweig Germany
2. Zoological InstituteTechnische Universität Braunschweig Braunschweig Germany
3. Institute of Earth Sciences Friedrich‐Schiller‐Universität Jena Germany
4. Department of Evolutionary Genetics Max Planck Institut für Evolutionsbiologie Plön Germany
5. Key Laboratory of Tibetan Environment Changes and Land Surface Processes Institute of Tibetan Plateau Research Chinese Academy of Sciences Beijing China