1. Spatial Epidemiology and Ecology Group, Department of ZoologyUniversity of OxfordOxford UK
2. Peter Medawar Building for Pathogen ResearchUniversity of Oxford, and Oxford NHIR BRCOxford UK
3. Institute of Health Metrics and EvaluationSeattle WA USA
4. Division of Infectious DiseasesSt Mary's Campus, Imperial CollegeLondon UK
5. Department of ZoologyUniversity of OxfordOxford UK