1. School of Basic MedicineQingdao University Qingdao 266071 P. R. China
2. CAS Key Laboratory of Tropical Marine Bio-resources and Ecology, Guangdong Key Laboratory of Marine Materia Medica, South China Sea Institute of OceanologyChinese Academy of Sciences 164 West Xingang Road Guangzhou 510301 P. R. China
3. College of OceanographyUniversity of Chinese Academy of Sciences Beijing 100049 P. R. China