1. Department of Medical Genetics, Cambridge Institute for Medical Research (CIMR), University of Cambridge, Cambridge CB2 0XY, United Kingdom
2. Department of Anatomy, Yonsei University College of Medicine, Seoul 03722, Korea
3. UK Dementia Research Institute, University of Cambridge, Cambridge CB2 0AH, United Kingdom
4. BK21 PLUS Project for Medical Science, and Brain Research Institute, Yonsei University College of Medicine, Seoul 03722, Korea