1. Unit of Plant Biochemistry, Scottish Crop Research Institute, Invergowrie, Dundee DD2 5DA, United Kingdom
2. Unit of Cell Biology, Scottish Crop Research Institute, Invergowrie, Dundee DD2 5DA, United Kingdom
3. Department of Evolutionary Biology, University of Parma, Viale delle Scienze, 43100 Parma, Italy
4. Unit of Gene Expression, Scottish Crop Research Institute, Invergowrie, Dundee DD2 5DA, United Kingdom