1. Faculty of Sciences of Gabès University, Zrig Gabès 6072 Tunisia
2. Arid Regions Institute Douz Road Kebili Tunisia
3. Department of Biology, College of Science Taif University PO Box 11099 Taif 21944 Saudi Arabia
4. CEFE, Univ Montpellier, CNRS EPHE‐PSL University, IRD Montpellier France