1. School of Biology; University of Leeds; Leeds LS2 9JT UK
2. Institute for Crop and Soil Science; Julius Kühn-Institute - Federal Research Centre for Cultivated Plants; Bundesallee 50 D-38116 Braunschweig Germany
3. Institute of Biodiversity; Thünen Institute - Federal Research Institute for Rural Areas, Fisheries and Forestry; Bundesallee 50 D-38116 Braunschweig Germany