1. ISEM Univ Montpellier, CNRS, IRD Montpellier France
2. Centre for Biodiversity Theory and Modelling, Theoretical and Experimental Ecology Station, CNRS and Paul Sabatier University Moulis France
3. PHIM Plant Health Institute, University of Montpellier, CIRAD, INRAE, Institut Agro, IRD Montpellier France
4. France Santa Fe Institute Santa Fe New Mexico USA