1. Marine Biology and Ecology Research Centre; University of Plymouth; Plymouth PL4 8AA UK
2. The Marine Biological Association of the United Kingdom (MBA); The Laboratory; Citadel Hill Plymouth PL1 2PB UK
3. Southern Seas Ecology Laboratories, Earth and Environmental Sciences; The University of Adelaide; Adelaide SA 5005 Australia
4. Australian Institute of Marine Science; PMB 3 Townsville QLD 4810 Australia