1. Disease Dynamics Unit, Department of Veterinary Medicine, University of Cambridge, Cambridge, UK
2. Department of Biology, Colorado State University, Fort Collins, CO, USA
3. Institute of Zoology, Zoological Society of London, London, UK
4. Animal Health and Veterinary Laboratories Agency, Weybridge, UK
5. Department of Biomedical Sciences, Colorado State University, Fort Collins, CO, USA
6. U.S. Geological Survey, Fort Collins Science Center, Fort Collins, CO, USA
7. Department of Ecology and Evolutionary Biology, University of Tennessee, Knoxville, TN, USA
8. National Center for Emerging and Zoonotic Infectious Diseases, Centers for Disease Control and Prevention, Atlanta, GA, USA