1. State Key Laboratory of Systematic and Evolutionary Botany Institute of Botany Chinese Academy of Sciences Beijing 100093 China
2. University of Chinese Academy of Sciences Beijing 100049 China
3. Jiangxi Province Key Laboratory of Watershed Ecosystem Change and Biodiversity Institute of Life Science and School of Life Sciences Nanchang University Nanchang 330031 China
4. Missouri Botanical Garden 4344 Shaw Blvd St Louis MO 63166‐0299 USA
5. Institut de Systématique, Evolution, Biodiversité (ISYEB) Muséum national d’Histoire naturelle CNRS Sorbonne Université Université des Antilles EPHE 57 rue Cuvier, CP39 Paris 75005 France