1. Oxford Regional Molecular Genetics Laboratory; Oxford University Hospitals NHS Trust; Oxford UK
2. Institute of Integrative Biology; ETH Zürich; 8092 Zürich Switzerland
3. School of Mathematics and Statistics; University of Sheffield; Sheffield S3 7RH UK
4. Institute of Evolutionary Biology; The University of Edinburgh; Edinburgh EH9 3JT UK
5. Biosciences; University of Exeter; Penryn Cornwall TR10 9EZ UK