1. School of Life Sciences; Nottingham NG7 2UH UK
2. National Institute of Environmental Health Sciences, National Institutes of Health; Department of Health and Human Services; Research Triangle Park NC 27709 USA
3. School of Physiology, Pharmacology and Neuroscience; University of Bristol; Bristol BS8 1TD UK
4. Neuroscience Division, Medical Education Institute, Ninewells Hospital and Medical School; University of Dundee; Dundee DD1 9SY UK
5. Centre for Integrative Physiology; University of Edinburgh; Edinburgh EH8 9XD UK