1. Department of Infection Biology; Institute of Infection and Global Health; School of Veterinary Science; University of Liverpool; Liverpool L3 5RF UK
2. CIGMR (Centre for Integrated Genomic Medical Research); Faculty of Medical and Human Sciences Manchester University; Manchester M13 9PT UK
3. Department of Musculoskeletal Biology; Institute of Ageing and Chronic Disease; University of Liverpool; Liverpool CH64 7TE UK