1. UCA, INRA, PIAF; 63000 Clermont-Ferrand France
2. UCA, EA 4678 CIDAM; 63000 Clermont-Ferrand France
3. Agriculture and Botany Department, Faculty of Agriculture; Menoufia University; Shebin El-Kom Egypt
4. Department of Forest Genetics and Plant Physiology; Swedish University of Agricultural Sciences; 901 83 Umeå Sweden