1. Department of Biology University of Utah Salt Lake City UT 84112‐0840 USA
2. Centro de Investigación de la Biodiversidad y Cambio Climático (BioCamb) e Ingeniería en Biodiversidad y Recursos Genéticos Facultad de Ciencias de Medio Ambiente Universidad Tecnológica Indoamérica Quito EC170103 Ecuador
3. Smithsonian Tropical Research Institute Box 0843‐03092 Balboa Ancón Republic of Panamá
4. School of Biological Sciences University of Tasmania Sandy Bay TAS 7001 Australia
5. Ashworth Labs Institute of Evolutionary Biology School of Biological Sciences University of Edinburgh Edinburgh EH9 3JY UK
6. Royal Botanic Garden Edinburgh Edinburgh EH3 5LR UK
7. School of GeoSciences University of Edinburgh Edinburgh EH9 3FF UK