1. GReD; Université Clermont Auvergne; CNRS; INSERM; BP 38 63001 Clermont-Ferrand France
2. The Sainsbury Laboratory; University of Cambridge; Cambridge CB2 1LR UK
3. Department of Plant Biology; Uppsala BioCenter; Swedish University of Agricultural Sciences; Uppsala 75007 Sweden
4. Department of Biostatistics and Bioinformatics; Rollins School of Public Health; Emory University; 1518 Clifton Road NE Atlanta GA 30322 USA