1. Biodiversity and Climate Research Centre (BiK-F) and Senckenberg Gesellschaft für Naturforschung; Senckenberganlage 25 60325 Frankfurt (Main) Germany
2. Department of Biological Sciences; Johann Wolfgang Goethe-Universität Frankfurt; Max-von-Laue-Straße 9 60438 Frankfurt (Main) Germany
3. Swiss Federal Research Institute for Forest, Snow and Landscape Research (WSL); Zürcherstrasse 111 8903 Birmensdorf Switzerland