1. Shanghai Chenshan Plant Science Research Center; Chinese Academy of Sciences; Shanghai Chenshan Botanical Garden; 3888 Chenhua Road Shanghai 201602 China
2. Landscape Architecture College; Beijing Forestry University; No. 35 Tsinghua East Road Haidian District Beijing 100083 China
3. Université Paris-Sud; Ecologie, Systématique et Evolution; UMR8079; 91405 Orsay France
4. CNRS; Ecologie; Systématique et Evolution; UMR8079; 91405 Orsay France
5. AgroParisTech; 91405 Orsay France