1. Marine Ecology; GEOMAR Helmholtz Centre of Ocean Research Kiel; Kiel 24105 Germany
2. Faculty of Mathematics and Natural Sciences; Christian-Albrechts-University; Kiel 24118 Germany
3. Department of Geography and Environmental Studies; University of Dodoma; P.O. Box 395 Dodoma 0105 Tanzania
4. Faculty of Science, School of Biology; Aristotle University of Thessaloniki; Thessaloniki 54124 Greece