1. AMAP, Univ Montpellier, CIRAD, CNRS, INRAE, IRD F‐34398 Montpellier France
2. Centre for African Ecology, School of Animal, Plant and Environmental Sciences University of the Witwatersrand Private Bag X3 WITS Johannesburg 2050 South Africa
3. Global Change Biology Group, Department of Botany and Zoology Stellenbosch University Private Bag X1 Matieland 7602 South Africa
4. Institute of Botany of the Czech Academy of Sciences v.v.i, Dukelská 135 Třeboň 379 01 Czech Republic
5. CNRS UMR7618 Institute of Ecology and Environmental Sciences Paris, Sorbonne University 4 Place Jussieu 75005 Paris France