1. MRC/Wits Developmental Pathways for Health Research Unit, Department of Paediatrics, School of Clinical Medicine, Faculty of Health Sciences; University of the Witwatersrand; Johannesburg South Africa
2. Division of Human Genetics, School of Pathology, Faculty of Health Sciences; University of the Witwatersrand and the National Health Laboratory Service; Johannesburg South Africa
3. Department of Obstetrics and Gynaecology, School of Clinical Medicine, Faculty of Health Sciences; University of the Witwatersrand; Johannesburg South Africa
4. Department of Paediatrics; University of Cambridge; Cambridge UK
5. Wellcome Trust-MRC Institute of Metabolic Science; Cambridge UK