1. Department of Biology; California State University; 18111 Nordhoff Street Northridge CA 91330-8303 USA
2. Center for Conservation and Evolutionary Genetics; Smithsonian Conservation Biology Institute; National Zoological Park; Washington DC USA
3. Department of Vertebrate Zoology; National Museum of Natural History; Smithsonian Institution; PO Box 37012 MRC 5503 Washington DC 20013-7012 USA
4. Department of Ecology & Evolutionary Biology; Brown University; Providence RI 02912-G USA