1. National Key Laboratory for Germplasm Innovation & Utilization of Horticultural Crops, Key Laboratory of Potato Biology and Biotechnology, Ministry of Agriculture and Rural Affairs Huazhong Agricultural University Wuhan 430070 China
2. Chongqing Key Laboratory of Biology and Genetic Breeding for Tuber and Root Crops, Engineering Research Center of South Upland Agriculture, Ministry of Education Southwest University Chongqing 400715 China
3. Department of Biology Hong Kong Baptist University Hong Kong Hong Kong Special Administrative Region (HKSAR) China
4. Plant Breeding Wageningen University and Research Droevendaalsesteeg 1 Wageningen 6708 PB The Netherlands