1. Department of Marine Science, Caspian Sea Basin Research Center University of Guilan Rasht Iran
2. Department of Biology, Faculty of Science University of Guilan Rasht Iran
3. International Sturgeon Research Institute Rasht Iran
4. Escuela Nacional de Ingeniería Pesquera, Lab. Bioingeniería Acuícola Universidad Autónoma de Nayarit Nayarit México