1. Max Planck Institute for Demographic Research Konrad Zuse Str.1 18057 Rostock Germany
2. School of Biological Sciences Centre for Biodiversity and Conservation Science The University of Queensland St Lucia Qld 4072 Australia
3. Max Planck Odense Center on the Biodemography of Aging Campusvej 55 DK‐5230 Odense M Denmark
4. Institute of Biology University of Southern Denmark Campusvej 55 DK‐5230 Odense M Denmark
5. School of Biomedical and Biological Plymouth University B420 Portland Square, Drake Circus Plymouth Devon PL4 8AA UK
6. Department of Mathematics and Computer Sciences IMADA University of Southern Denmark Campusvej 55 DK‐5230 Odense M Denmark