1. Centre for Regenerative Medicine University of Edinburgh Edinburgh UK
2. Division of Molecular and Clinical Medicine Ninewells Hospital and Medical School University of Dundee Dundee UK
3. NHS TaysideNinewells Hospital and Medical School Dundee UK
4. Division of Gastroenterology and Hepatology University of Michigan Ann Arbor Michigan USA
5. Gastroenterology Section VA Ann Arbor Healthcare System Ann Arbor Michigan USA