1. State Key Laboratory of Crop Stress Biology for Arid Areas (Northwest A&F University) Yangling Shaanxi 712100 China
2. College of Plant Protection Northwest A&F University Yangling Shaanxi 712100 China
3. INRA UMR 1349 IGEPP Institut de Génétique, Environnement et Protection des Plantes Domaine de la Motte Le Rheu Cedex 35653 France