1. University of Guam Marine Laboratory; UOG Station Mangilao GU 96923 Guam USA
2. Ocean Associates, contracted to NOAA Pacific Island Fisheries Science Center; Coral Reef Ecosystem Division, 1125B Ala Moana Boulevard; Honolulu HI 96814 USA
3. Institut de Recherche pour le Développement; Hawaii Institute of Marine Biology; Kaneohe HI 96744 USA