1. Tianjin Key Laboratory of Conservation and Utilization of Animal Diversity, College of Life Sciences Tianjin Normal University Tianjin China
2. Department of Fisheries, Wildlife and Conservation Biology University of Minnesota St. Paul Minnesota USA
3. Ministry of Education Key Laboratory for Biodiversity Science and Engineering, College of Life Sciences Beijing Normal University Beijing China
4. National Forestry and Grassland Administration Key Laboratory for Conservation Ecology of Northeast Tiger and Leopard Beijing China