1. Department of Organismic and Evolutionary Biology; Harvard University; 26 Oxford Street Cambridge MA 02138 USA
2. Department of Biological Sciences; George Washington University; Lisner Hall Room 340 2023 G Street, NW Washington DC 20052 USA
3. Laboratory of Insect Social Evolution; The Rockefeller University; 1230 York Avenue New York NY 10065 USA
4. Instituto de Biologia; Universidade Federal de Uberlândia (UFU); Caixa Postal 593 38400-902 Uberlândia MG Brazil
5. Department of Ecology and Evolutionary Biology; University of Toronto; 25 Harbord Street Toronto ON M5S3G5 Canada