1. Key Laboratory of Zoological Systematics and Evolution Institute of Zoology Chinese Academy of Sciences Beijing China
2. Severtsov Institute of Ecology and Evolution RAS Moscow Russia
3. CAS Key Laboratory of Mountain Ecological Restoration and Bioresource Utilization & Ecological Restoration and Biodiversity Conservation Key Laboratory of Sichuan Province, Chengdu Institute of Biology Chinese Academy of Sciences Sichuan China
4. Xinjiang Key Laboratory of Biological Resources and Genetic Engineering, College of Life Science and Technology Xinjiang University Urumqi China
5. The Key Laboratory of Zoological Systematics and Application, School of Life Science, Institute of Life Science and Green Development Hebei University Baoding China