1. Department of Ecology and Evolutionary Biology; Princeton University; 106A Guyot Hall Princeton NJ 08544 USA
2. Department of Ecology, Evolution and Natural Resources & Institute of Marine and Coastal Sciences; Rutgers University; 14 College Farm Rd. New Brunswick NJ 08901 USA
3. Department of Biology; Stanford University; Hopkins Marine Station 120 Ocean View Blvd. Pacific Grove CA 93950 USA