1. CAS Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany Chinese Academy of Sciences Kunming 650201 China
2. Department of Economic Plants and Biotechnology, Yunnan Key Laboratory for Wild Plant Resources, Kunming Institute of Botany Chinese Academy of Sciences Kunming 650201 China
3. University of Chinese Academy of Sciences Beijing 100049 China
4. Global Biodiversity Conservancy 144/124 Moo3, Soi Bua Thong, Bangsalae, Sattahip Chonburi 20250 Thailand
5. Key Laboratory of Southern Subtropical Plant Diversity, Fairy Lake Botanical Garden Shenzhen & Chinese Academy of Sciences Shenzhen 518004 China
6. Department of Earth and Environmental Sciences Indiana University‐Indianapolis Indianapolis 46202 IN USA