1. Department of Microbiology and Molecular Genetics, Briggs Hall, One Shields Avenue University of California Davis CA USA
2. MRC Human Genetics Unit, Institute of Genetics and Molecular Medicine University of Edinburgh Crewe Road South, Edinburgh, EH42XU, Scotland UK
3. Institute for Global Food Security, School of Biological Sciences Queen's University Belfast 19 Chlorine Gardens, Belfast, BT9 5DL Northern Ireland UK